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Urban Food Security. Therefore, formal comparison between both taxa is for now precluded. Line drawings of the holotype tympanic bulla MCZ of Monotherium? Calvert Formation at Richmond, Virginia in ventral view. After figures from Ray [ 15 ]. Broken and obliterated parts are indicated in grey. Ray Prostitutes Orekhovo-Zuyevo 15 ] provided evidence supporting Wyman's statement. Type horizon. However, the Calvert Formation spans across the entire early Miocene and into the late middle Miocene ca At the Calvert Cliffs in Maryland, the oldest published record phocid fossils come from zone 10 of the Calvert Formation [ 64 ] which is dated to the early middle Miocene [ 65 ]; and references therein.

This renders a pre-middle Miocene age for MCZ less likely. Originally presented as Phoca wymaniRay [ 15 ] tentatively placed the original material of the species in Monotherium? The fossil record of M. Hence, the diagnostic value of fossil phocid ear regions is undermined. Additionally, a study of tympanic bullae of elephant seals, Miroungahas shown that intraspecific variation is prominent [ 52 ]. Because other specimens attributed to M. Moreover, the fossil record of referred specimens of M.

The lack Prostitutes Orekhovo-Zuyevo humeri in the fossil record of M. The holotype ear region of M. Prostitutes Orekhovo-Zuyevo characteristics clearly support the identification of MCZ as a monachine e. However, because the genus Monotherium is restricted to its type species, M. The other specimens tentatively referred to M. Muizon Prostitutes Orekhovo-Zuyevo 42 ] already noted the value of the shape of the trochlear notch of the ulna as a means to distinguish Phocidae to the generic Prostitutes Orekhovo-Zuyevo, also indicating similarities between the ulnae of Phocidae cf.

Contrastingly, we deem it inappropriate to identify the mandible and the partial tibia and fibula of USNM beyond the subfamily level. Ray [ 15 ] stated that the specimens came from one single block and are probably of one individual, but he implicitly expressed his doubt.

Indeed, their association can be considered Prostitutes Orekhovo-Zuyevo in the absence of other Prostitutes Orekhovo-Zuyevo parts of the skeleton. Pending the discovery of more complete fossil specimens of Monotherium? Apart from M. However, Dewaele et al. And Prostitutes Orekhovo-Zuyevo, new research by Dewaele et Prostitutes Orekhovo-Zuyevo. Thus, the holotype of M. Therefore, we consider M.

Despite being uncomparable to Monotherium delogniiwe provisionally retain the genus name Monotherium with a question mark.

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It is unknown whether M. It is likely that the holotype will eventually be designated into another genus, but this cannot be ascertained based on the current fossil record. Contrastingly, Monotherium gaudini is given a new genus name in this study, based on the argumentation that the systematic comparison and phylogenetic analysis see above and below Prostitutes Orekhovo-Zuyevo the more complete type material places it as a Prostitutes Orekhovo-Zuyevo monachine, while the phylogenetic affinities of the Prostitutes Orekhovo-Zuyevo of M.

The current genus name Monotherium is preferred as placeholder Prostitutes Orekhovo-Zuyevo the genus name Phoca, which was used prior to Ray's redescription [ 15 ].

The name Phoca is currently restricted to Phoca largha and Phoca vitulina : Prostitutes Orekhovo-Zuyevo phocine seals that show no affinities with M. Identification as a phocid seal supported by the large development of the deltopectoral crest on the humerus. Identified as a phocine Prostitutes Orekhovo-Zuyevo on the presence of an entepicondylar foramen also in Homiphoca capensis and the overall slenderness. Differs from most Phocinae by having a very strongly reduced Prostitutes Orekhovo-Zuyevo neck also in Histriophoca fasciataLeptophoca proxima and Pagophilus Prostitutes Orekhovo-Zuyevo.

Differs from all Phocidae in the following unique combination of characteristics: lesser tubercle slightly below the level of the humeral head also in Devinophoca emryiLe. Here, the term is used Prostitutes Orekhovo-Zuyevo reference to the Mare Frisicumthe Latin name for the North Sea, alluding to the geographical origin of the two species of the genus listed here.

Although Frisiphoca shares characteristics with both Monachinae and Phocinae see diagnosisthe presence of an entepicondylar foramen is Prostitutes Orekhovo-Zuyevo as a characteristic uniting Frisiphoca with Phocinae.

Unfortunately, this is not reflected in the phylogenetic analysis due to the poor scoring of the fragmentary fossil record of the genus. Corresponding labelled drawings of right humerus IRSNB M of Frisiphoca aberratum in posterior imedial janterior k and lateral l view; and lectotype right humerus IRSNB M of Frisiphoca affine in posterior mmedial nanterior o and lateral p view. Daily update on what's on in gay Ekeren. Finale Gala Mr Gay Belgium And not to mention the beers, you can have a look Prostitutes Orekhovo-Zuyevo Belgium's beer culture with his guidance.

Read Prostitutes Orekhovo-Zuyevo. Date of experience: August. Scale bar equals 10 cm. Van Beneden nor any subsequent author assigned a type specimen to Frisiphoca [ Monotherium ] aberratum. In the absence of more completely preserved material, we consider the humerus figured by Van Beneden [ 17 ]; pl. Third section at Borgerhout see original label with specimenAntwerp, Belgium. However, it should be noted that this type locality is derived Prostitutes Orekhovo-Zuyevo the original labels associated to the specimen.

Prostitutes Orekhovo-Zuyevo his original publications Van Beneden [ 1617 ] did not discuss the geographical provenance of individual specimens of the original fossil record of Frisiphoca aberratum. However, as mentioned above, the Diestian is currently considered an obsolete term and should not be used any more [ Prostitutes Orekhovo-Zuyevo ]. Different authors assign different ages Prostitutes Orekhovo-Zuyevo stratigraphic intervals to the Diestian see [ 19 ]: table 1but in general it is considered that the Diestian is roughly equivalent to the Deurne Sands Member of the Diest Formation.

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In a more detailed description of the Neogene stratigraphy of the Antwerp area, Mourlon [ 68 ] mentions the occurrence of Frisiphoca aberratum and possibly also Frisiphoca [ Monotherium ] affine and Monotherium delognii in one of the strata he described. This unnamed stratum, composed of greenish glauconiferous sands overlays—also unnamed—darker green and black sands [ 68 ]. Extrapolating this to the current knowledge on the Prostitutes Orekhovo-Zuyevo of the Antwerp area, these greenish glauconiferous sands most likely represent the Deurne Sands Member of the Diest Formation, while the underlying darker green sands and black sands represent the Antwerpen Sands Member of the Berchem Formation [ 69 ].

Therefore, it can be assumed that F. Selection of cranial measurements of Noriphoca gaudini redrawn from Guiscardi [ 7 Prostitutes Orekhovo-Zuyevo originally for Phoca gaudini. Medium-sized phocine, comparable in size to the harbour seal, Phoca vitulina. Differs from all Phocidae, including Frisiphoca affineby the strong posteroproximal Prostitutes Orekhovo-Zuyevo of the humeral Prostitutes Orekhovo-Zuyevo, and differs from Phocinae, including F.

Differs Prostitutes Orekhovo-Zuyevo from F. Given the overall rarity of such bones in the fossil record, they cannot be compared with other extinct phocid taxa from the southern North Sea basin, and the lectotype humerus has been selected as the Prostitutes Orekhovo-Zuyevo specimen of F. A sediment sample associated with the vertebrae IRSNB M was analysed biostratigraphically with dinoflagellate cysts for our study.

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The humerus IRSNB M is overall well preserved, only missing part of the deltopectoral crest and portions of the distal epiphysis. The bone is straight and moderately slender. Ray [ Prostitutes Orekhovo-Zuyevo ] already noted that Prostitutes Orekhovo-Zuyevo humerus is relatively straight in some early Phocidae, such as Leptophoca proximaas well as early stem pinnipedimorphs and terrestrial carnivorans, while most extinct and recent Phocidae have a more strongly curved Prostitutes Orekhovo-Zuyevo diaphysis.

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The humerus of F. Measurements with an asterisk are estimations. Measurements in mm. Sources provided for measurements retrieved from the literature. Measurements by Koretsky [ 2 ] represent averages of multiple specimens. The humeral head is small and strongly hemispherical. As in Acrophoca longirostrisHydrurga leptonyx and Otariidae, the head faces relatively proximally contra [ 42 ]relatively more posteroproximal than in other Phocidae, including F.

Unlike other Phocinae, Prostitutes Orekhovo-Zuyevo the extinct Leptophoca proxima [ 10 ], and the extant Prostitutes Orekhovo-Zuyevo fasciata and Pagophilus groenlandicusthe neck is poorly developed in the species of Frisiphoca. Although incompletely preserved, it appears that the deltoid tuberosity must have been located approximately halfway the length of the bone. The bicipital groove bears a small but noticeable transverse bar, which is also observed in the Monachinae Lobodon carcinophoca, Monachus spp.

The lesser and greater tubercles reach slightly below Prostitutes Orekhovo-Zuyevo level as the head contra [ 42 ]; for the greater tubercle. Among Phocidae, most extant taxa except Monachus have a strongly developed Prostitutes Orekhovo-Zuyevo tubercle and a small greater tubercle. While this condition is variable Prostitutes Orekhovo-Zuyevo extant taxa, many early phocid taxa bear a relatively little-developed lesser tubercle e.

Leptophoca proximaNanophoca vitulinoides. Geologically younger extinct Phocidae e. Prostitutes Orekhovo-Zuyevo capensisPliophoca etrusca tend to have a lesser tubercle that shows a degree of development intermediate between early Prostitutes Orekhovo-Zuyevo Phocidae and extant Phocidae [ 9 ]. On the lateral surface of the greater tubercle, the insertion area for the infraspinatus and supraspinatus muscles is well outlined. The deltopectoral crest is overall slender and tapers smoothly towards Prostitutes Orekhovo-Zuyevo coronoid Prostitutes Orekhovo-Zuyevo, distally, as is typical for many extinct monachines and phocines see genus diagnosis [ 1042 ].

Prostitutes Orekhovo-Zuyevo distal to the humeral head and lesser tubercle, on the posterior surface of the diaphysis, there is a prominent fossa for the origin of the triceps brachii muscles. An entepicondylar foramen is present, which is a characteristic shared with other Phocinae; among Monachinae, only Homiphoca capensis has Prostitutes Orekhovo-Zuyevo entepicondylar foramen [ 4258 ].

The supinator crest is reduced, as in Monachinae, but contrasting to other Phocinae, including F. The medial epicondyle is broad and flaring, as in F. The lateral margin of this supinator crest is rugose, providing an origin area for powerful manual extensor muscles. The medial epicondyle is broad at the distal extremity of the supinator crest and medial to the medial condyle. The olecranon fossa and coronoid fossa are strongly reduced, these regions being almost completely flat.

When compared to other Phocidae except Homiphocaat the distal epiphysis the ulnar articular facet of the trochlea is prominent in relation to the radial capitulum.

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This contrasts with F. Overall, the humerus of F. Prostitutes Orekhovo-Zuyevo poorly developed supinator crest is considered a plesiomorphic character retained in Monachinae, while the presence of an entepicondylar foramen is regarded as a plesiomorphic character retained in Phocinae [ 858 Prostitutes Orekhovo-Zuyevo.

Frisiphoca affine? Calvert Formation at Richmond, Virginia in medial b and anterior c view. USNM was considered? Referred specimen. In Prostitutes Orekhovo-Zuyevo redescription of Monotherium? Monotherium aberratum [ 15 ]: figs 8—11, subset 1. However, given the poor state of preservation of the specimen, we deem it best to consider the specimen Phocidae cf.

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Just distal to the humeral head and lesser tubercle, on the posterior surface of the diaphysis, there is a prominent fossa for the origin of the triceps brachii muscles.

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